Shkodra Region
Here's just a small sampling of the natural treasures of Shkodra Region.
Shkodra region is situated in the North-West part of Albania. It has a surface of 158.800 ha. There are five protected areas in Shkodra region
Shkodra Lake - Managed Natural Reserve
Buna River, Velipoje - Protected Landscape
Albanian Alps - National Park
Munella - Natural Park
Shala Valley - Natural Park
There are also 113 Natural Monuments in Shkodra region.
Shkodra Lake
Managed Natural Reserve, Ramsar Site (2005), Surface 24.050 ha
Shiroka, Shkodra Lake © Alketa Misja Photography
Shkodra Lake is located in south of European Continent, in the west of the Balkan Peninsula, in southwestern side of the state border between Albania and Montenegro. Shkodra Lake is included in the water system of the Adriatic Sea basin and it is right next to it.
The lake was formed by geotectonic developments of a very wide region and also powerful karst developments. The base contours of Southeast Dinaric, Zeta-Shkodra lowland and Shkodra Lake are consistency of endogenous effects during Mesozoic and partially Cenozoic. Downward movement of the block on which the lake is located started in the end of Paleogene and beginning of Neogene. These phenomena are in progress also today, which is important in forecasting the long-term developments of the lake and the region.
Shkodra Lake is the biggest lake of the Balkans with a surface area of approximately 360 - 540 km². Watershed Basin of Shkodra Lake has a considerable area surface of 5490 km². Shkodra Lake is a plain lake, but it is supplied with water from the height with more rainfall, with 2500 - 3100 mm rain per year; this is another factor of the stability of water flows in the lake. The water comes to the lake through rivers, streams, different sources, etc. Around 62% of the water comes from Moraça River. The emptying of the lake is realized through Buna River.
In the lake water regime, an important role plays the highly variable Drini River regime. Drini River flows into Buna River in about 1.5 km from the outlet of the lake, through the main tributary Drinasa. Shkodra Lake functions as a perfect hydrological stabilizer. When Drini brings some water, the lake pours water freely in Buna River. On the contrary, when Drini River brings too much water, the normal flow of Buna River is obstructed, so the flow of the lake water into Buna River is prevented, as a result the lake water increase its level, surface and its volume.
The lake is at the end of Zeta-Shkodra lowland. 165 km2 from the lake surface, which belongs to 44 % of its minimal surface, has the bottom under the sea level.
Hydrothermal regime of Shkodra Lake is interesting. The average temperatures from January to August change with about 20°C. Very rare the temperature has dropped under 0°C. This temperature regime characterizes the lake as subtropical. The lake has more water in rainy seasons and less during summer.
Predominance of clear waters, oligotrophic, with low biomass of phytoplankton, is identifying characteristic of the lake. Feedbacks that keep in normal parameters lake water quality are: nutrient content in the water versus humus production in the wetland; nutrient content in the water versus the production of coastal forest habitats; the structures of food chain that transfer phosphorus from littoral in pelagic versus biogeochemical mechanisms that inhibit recycling of phosphorus by sediment.
The high nutrient content in the lake sediment is expression of the normal development of the lake. This is related to decay and mineralization of macrophyte biomass, its distribution in the entire lake, and also the inhibit organisms of the phosphorus recycling by the sediment in the water.
Shkodra Lake consists of three habitat systems: the lacustrine system, that include limnetic and littoral habitats; palustrin system with differentiating habitats regarding vegetation, and also the presence or absence of water; riverine system where are included permanently flooded habitats and those often flooded. According to the seasons Shkodra Lake develops from a habitat complex of lake system in a habitat complex of lake-swamp system, with a multiply number of ecological niches. Littoral subsystem area of the lake, during the warm seasons moved towards depth, giving way the swamp habitats system.
The abundance of macrophytes with roots is identifying characteristic of Shkodra Lake. Macrophytes are the most characteristic feature of the littoral and in a sense also of the entire Shkodra Lake. The lake macrophytes develop from the coast towards depth, starting from emergent associations, stereotypically dominated by reed Phragmites australis, to associations with floating leaves, dominated by yellow water lily Nuphar luteum, white water lily Nymphaea alba and water chestnut Trapa natans, continuing in those submerge, dominated by eurasian water milfoil Myriophyllum spicatum, coontail Ceratophyllum demersum, potamogeton Potamogeton and white water crowfoot Ranunculus aquatilis and that end to the deepest associations with domination of straight vallisneria Vallisneria spiralis, spiny water nymph Najas marina and Charophyta. In Shkodra Lake are known 240 water plant species, from which 62 are true aquatic species. 21 species are trees or shrubs. Until now in Shkodra Lake are known 60 plant associations.
The list of fishes (Actinopterygii) of Shkodra Lake and affluent waters contain 61 species. Cyprinidae family has 25 species, which consider as a high percentage and this fact characterizes the lake and its ichthyofauna as cyprinid. The waters around the lake and its ichthyofauna are salmonid. In the lake and its affluent are found 9 species of the family Salmonidae (17.3 %). The fishing in Shkodra Lake base in three groups of species. An important value for fishing have two native species, bleak (Alburnus scoranza) and carp (Cyprinus carpio). In the other group are included species migration to the sea through Buna River, where the most important are twait shad (Alosa fallax), European eel (Anguilla anguilla) and mullets (Mugil cephalus and Liza ramado). The third group includes introduced species, as carassius (Carassius gibelio) and European perch (Perca fluviatilis).
The biology of Shkodra Lake fishes and fishing has intriguing features. The considerable fish stocks are caught in crypto depressions or in some special places, where it pass the winter. The fish reproduction is generally done along the coast. The decrease of the market fish populations is related with human activity, mainly with illegal fishing.
Shkodra Lake stands out for its diversity, capacity and great ornithological role in regional level. In Shkodra Lake and its watershed basin have been recorded 283 bird species, from which 168 are of water habitats. From this number, more than half are nesting birds. In Shkodra Lake is found and nest the global endangered species of Dalmatian pelican Pelecanus crispus. Shkodra Lake meets the Ramsar criterion for the number of nesting pairs of whiskered tern Chlidonias hybridus and pygmy cormorant Phalacrocorax pygmeus. More than 2/3 of the total species of water habitats is migratory birds. Shkodra Lake is through the most important wetlands in Europe for birds wintering, with a capacity of 250.000 individuals, meeting so the Ramsar criterions for the total number of individuals. The largest number of the individuals comprised by Eurasian coot (Fulica atra), cormorants (Phalacrocorax), grebes (Podiceps), some ducks (Anatidae), gulls (Larus), etc. The decrease of species and individuals number is related to negative human impact, disruption of the water regime, hunting and different concerns.
Shkodra Lake is diagnosed of 8th category in the gradient with 10 naturalness categories. The high naturalness is one of Shkodra Lake ecosystem qualities. Generally there is no addition from outside of matter in the lake, not physical alternation of the geomorphology or the availability of physical elements. Dynamics in the lake are natural. The level of habitats fragmentation considered moderate.
Fishing in Shkodra Lake is considered large. In the fish community there is the dominance of native species and the presence of exotic species, but with not a big impact.
In the lake coasts are collected sand and other materials. The artificial elements, as commercial buildings, quaysides and roads are not dominant in the lake landscape. The waste waters, items originating from waste, agriculture chemicals we think they can be processed and do not go beyond resilience.
The phenomenon of endemism if more pronounced in some animal groups of the lake. 10 mollusk species, of Gastropoda class, in the lake and downstream flows of its tributaries show the phenomenon of endemism, or are new for the science. Also 9 crab species Amphipoda that make up about half of the total number of species of this group in this lake are endemic.
A special interest present 20 fish species or 38.5% of the total number of fish species, which show the phenomenon of endemism. From them 5 are endemic of Shkodra Lake, 4 species are endemic of water ecosystems in Albania, 5 species are endemic of hydrological system Shkodra Lake – Drini River – Ohrid Lake, 4 species are endemic of Western Balkan, and two species are endemic of the lake with fluctuations towards the north.
Balkan frog Pelophylax kurtmuelleri is sub endemic for Albania, while Albanian frog Pelophylax shqiperica endemic of Shkodra Lake and Western lowland of Albania.
In trees, birds and many other groups, endemism is presented at the subspecies level.
High resilience is a characteristic of Shkodra Lake ecosystem. The high resilience is maintained by slow variables, as: long-term climatic factors, trophic relations, habitat structures, nutrients production, nutrient level in the sediment, etc.
Macrophytes with roots are a slow variable factor that promotes resilience. Also the feedbacks that maintain the water quality in normal parameters are held by resilience; otherwise the situation would degenerate in unwanted surprises. Damage to the functional group of fishes and organic polluters discharge may bring the loss of the ecological resilience.
The high stability is another characteristic of Shkodra Lake. This lake is identified as a shallow lake, with clear water and high abundance of macrophytes with roots. The high resilience provides high stability. The factors that influence for stability and high resilience of the lake are: high fluctuation of the nutrients in the lake, food chains with relatively with few links, large number of species, especially fishes that transfer phosphorous from the littoral to the pelagial.
Three feedbacks are main factors that maintain lake water quality at the normal parameters.
Macrophytes are considered key factor of stability. They produce nutrients, serve as links in the food chains, are the most important habitats for animals, are inhibiting factors of the nutrients recycling (especially phosphorus) from the sediment to the water, etc.
Shkodra Lake has a high interspecific diversity where could be mentioned: water plants 240 species, water mollusks in the basin 91 species, insects in the basin 6000 species, fish 61 species, amphibians in the basin 15 species, reptilians in the basin 28 species, water reptilians 4 species, birds in the basin 283 species, water birds in the lake and river 168 species, mammalians in the basin 57 species and water mammals 3 species.
Buna River
Protected landscape, Ramsar Site (2005). Surface 21.679 ha
Buna River © Alketa Misja Photography
Buna River (Serb. Bojana, Lat. Barbana) is 44 km long, 0.35 – 0.5 km wide and 2 - 4 m deep. The river begins in southernmost extremity of Shkodra Lake, it is the only emissary of the lake and flow to the Adriatic Sea forming a natural delta. The state border with Montenegro runs along the Buna River from the villages Samrisht in Albania and Gorice in Montenegro, until it flows to the sea.
Buna River Basin consists of three main types of rocks: Triassic, upper Crust and Eocene. Two types of geological base are distinguished: permeable rocks – limestone, and also no permeable rocks – flysh and quaternary sediments. The permeability of the rocks and erodibility affect directly a number of environmental characteristics, as the morphology of land shaft and habitat, hydrogeological and hydrological features, hydrochemistry of the running water and its temperature, factors which affect the biota.
In 1822, Drini and Buna were separated. Drini flowed directly at Adriatic Sea, near Lezha, about 25 km to the east of the actual Buna Delta. In 1867, a branch called Drinasa passed a part of the water from Drini in Buna River. In 1914 Drini discharged most of the water in Buna River through Drinasa. This situation develops also today. Buna River flows out from Shkodra Lake and only 1.5 km from the exit join the Drinasa water, the biggest tributary branch of Drini River.
The bottom of Buna River bed is some meters under the sea level. This makes sea water to penetrate along its bed’s bottom.
Buna River has a permanent river regime that is characterized by continuous water flow during the entire year in the river channel.
Buna River takes from the lake approximately 320 m³/sec water and from the Drini River branch takes 360 m³/sec, so in total 680 m³/sec. The average discharge of Buna River is 21.2 km3 /year, being so the third river in European Mediterranean, after Rhone and Po rivers.
Buna River can be considered as part of the unique hydrological complex, composed of Moraca River, Drini River, Buna River and big cross border lakes of Shkodra, Ohrid and Prespa. Buna River represents the final plain segment, genuine potamic of this complex.
The water basin where Buna River collects its water from is 20,585 km². The average hight of this basin is 909 m over the sea level. Buna River collects water from a hydrographic network extending in five countries: Albania, Montenegro, Kosovo, North Macedonia and Greece. Buna River collects water also from the streams, especially them of Ana Mali (in Shkodra), where the longest one is Milla Stream, 25 km.
The alluvial sediments are mainly gravel, sand and clay. The large materials deposit in the bottom of the river bed, especially in the upstream, while the others are transported in suspension, or in solution. The transport of the sediments along the river occurs mostly in winter or autumn, when the river water level reaches the highest level.
The estuary of Buna River is a coastal body of salty water, where two tributaries of the river flow and which has a free connection with open sea. The estuary represents a transition area between river and marine environments. So, the estuary is under two impacts: marine, which means waves and salt water flow, and also riverine, which means the flow of fresh water and sediments. The estuary filled by sea water during the ebb and with river water during the tide. The sea water and fresh water from the river provide high level of nutrients in the water and sediments; therefore the estuary of Buna River is the most productive natural habitat in the entire complex of the river.
Buna Delta represents a sediment surface, created where the river joins the sea, but in the opposite direction with that of the estuary. Its formation is favored by sea waves, and also large flows of sediments from Drini River. The actual natural delta is formation from the actual Buna River and the sea. After the construction of the hydropower artificial lakes, the sediment flows have decreased till 30 % and so the front part of the delta has begun to be destroyed.
The regime of water and sediments flow of Buna River is messed up after the construction of the dams and lakes of the hydropowers in Drini River. The forced discharges cause the disorder of the water regime and often floods. The Drini sediment flow quantity has a large decrease. Erosion is found in the lands near the river. There is habitat loss or damaged, surface denaturation because of the constructions, services, etc. The increase of the human presence, especially last decades because of the beach has led to the decrease of the birds population, especially water birds.
In the entire complex of the waters along Buna River, from Shkodra Lake till Adriatic Sea, are defined 29 habitat classes. Buna Delta is the most important wetlands system along Adriatic coast and one of the best preserved in Mediterranean and Europe. The river estuary stands out as exemplary, with highest naturality in Mediterranean.
Buna River stands out for its large water and nutrient flows. For these reasons are developed phytoplankton (with main groups Chlorophyta, Diatome and Cyanophyta), micro fauna (especially crabs, worms Rotifera and Protozoa), zoobenthos (dominated by crabs Crustacea, worms Oligochaeta, mollusks Mollusca, as Balcanic river snail Viviparus mamillatus, Bithynia and Holandriana, small river mussel Unio crassus, small tooth river mussel Microcondylaea compressa; insects Insecta, especially larvae Ephemeroptera, Plecoptera, Trichoptera, Diptera); macrophyte vegetation, cyprinid fishes, etc. We can distinguish salty waters of the estuary as very rich with phytoplankton and zooplankton, invertebrates (worms, mollusks and srabs), fishes and water birds.
Plants are abundant in rivers with low flow, as well as in Buna River. The flora of the area, related to water and humidity, contains over 170 species. Buna River macrophyte vegetation is located mainly in the river coasts from the exit from the lake till at the joint point with Drini River, and also at river mouth. It includes three plant association groups: submerge, floating and emerged.
Fauna is very rich in all its groups. Fishes and birds have high number of species. Amphibians are quite widespread in Buna river waters and around it. Especially are found tailless amphibians, through which the Albanian water frog Pelophylax shqiperica, endemic of low coastal Albania, Ulcinj lowland and Shkodra Lake. In this area are found 18 reptile species, from which 1/3 are endemic or subendemic taxa of the Balkans. Grass snake Natrix natrix, water snake Natrix tessellata and European pond turtle Emys orbicularis are very common in the waters. From the water mammals, otter Lutra lutra is found several times in Buna River. In Buna area there are quite a few mammal species, but most interesting is the finding of the footprints of brown bear Ursus arctos at the sand of Rrjolli beach.
The largest parts of the fishes, with about 30 species that are found in Buna River are the same of Shkodra Lake and mostly of them belong to the Cyprinidae family. These two indicators give to Buna River ichthyofauna a subtropical character like a lake one. About half of this number is endemic species of this hydrological system, or its parts. Such species are bleak Alburnus scoranza, western Balkan barbel Barbus rebeli, Albanian roach Pachychilon pictum, two species of (common) roachs Rutilus, Shkodra rudd Scardinius knezevici, Shkodra goby Pomatoschistus montenegrensis, Drini trout Salmo farioides, etc.
At the Buna mouth till 40 m deep into the sea, are found 66 fish species, especially benthic. More frequent are red mullet Mullus barbatus, European hake Merluccuis merluccius, picarel Spicara smaris, spotted flounder Citharus linguatula, large-scaled gurnard Lepidotrigla cavillone, etc. There are known 8 fish species that realize biological migration from Adriatic Sea, through Buna River, into Shkodra Lake and Drini River, and also vice versa. Flathead grey mullet Mugil cephalus, thin lip grey mullet Liza ramada, European eel Anguilla anguilla and twaite shad Alosa fallax are actually important species for the fish market in Shkodra. Whereas migratory species European sea sturgeon Acipenser sturio, Adriatic sturgeon Acipenser naccarii, seabass Dicentrarcus labrax and European flounder Platichthys flesus, last decades are rare too much or from a long time have not been seen. For all these species Buna River serves as a migration corridor from an ecosystem to the other one.
Diversity and abundance of fish in the Buna mouth area, and also 8 migratory fish species of Adriatic Sea, Buna River and Shkodra Lake ecosystems, represent outstanding cases of the integrated biological potentials of the ecosystems complex.
On the surface Ulcinj – Viluni Lagoon – Shkoder are recorded 237 bird species, from which 59 are water bird species. In the cold seasons of 2003-2020 are counted in total 16.964 - 25.328 water birds, which belong to 45 - 57 species. Among them visit these habitats Dalmatian pelican Pelecanus crispus, flamingo Phoenicopterus roseus, whooper swan Cygnus cygnus, etc. Mostly of this number was found in the Ulcinj saltpans and Zogaj Lake in Ulcinj, less in Viluni Lagoon, Shas Lake, and the smallest part in Velipoja Reserve, Domni marshland, Murtemsa Lake, etc. Most of the individuals belong to the species: Eurasian coot Fulica atra, black-headed gull Larus ridibundus, ducks Anatidae (single out velvet scoter Melanitta fusca), great cormorant Phalacrocorax carbo, black-throated loon Gavia arctica, dunlin Calidris alpina, etc. Over 40 % of wintering species are nesting birds or probably nesting. The other part of 60 % wintering here, but migrate for nesting in spring.
The estuary is one of the most interesting areas for water birds, especially wintering birds, mainly Phalacrocoracidae, Scolopacidae and Anatidae; related to it also Prodelta, very developed, which represents the part of the delta inside the sea and which has a relatively low rate of deposits.
Buna River and waters along it constitute a corridor for migratory birds along Adriatic coast.
Bottlenose dolphin Tursiops truncatus is common in sea waters of Albania and during the entire year prefers sea coasts and river mouths. In hot seasons are found and photographed these dolphins inside Buna River, at Velipoja village level, near Paratuk Island, near Shasi Lake, and an individual in Daragjat, 8-9 km from Shkodra Lake. For these reasons, Buna River can be considers as a dolphin’s river.
The sea turtle Caretta caretta deposits eggs into the sand of sea coasts, found also in Ada Island. This turtle enter also in short distances into the rivers, but an individual of 56 kg and 87 cm long, one decade before entered Buna River until Muriqan village. This case is very interesting for the science and constitutes an ecological value more for the hydrological complex Adriatic Sea – Buna River
Albanian Alps
National Park (2022). Surface 82.845 ha
Valbona Valley © Alketa Misja Photography
The Alps form a mountainous area in the northern part of Albania. They represent the highest and most majestic physiographic-geographic region, and their borders are quite clear. They extend to the north of the Drin River, west of the lower course of the Valbona River (a branch of the Drin). Their total area reaches 2,020 km2, occupying about 8% of the country's surface. The vegetation (plant cover) is understood as a complex mosaic of plant communities, which vary more or less continuously in space, in accordance with changes in environmental conditions (climate, relief, geological composition, soil, hydrological conditions). Thus, the vegetation of the Alps has undergone changes during its historical development.
In this territory, there are species of mammals and birds that are quite rare for the Balkan region and beyond. Some of them are relics, endemic, threatened with extinction, and for this reason, they are listed in international red books. Among the mammal species are Lynx lynx (Eurasian lynx), Ursus arctos (Brown bear), Canis lupus (Gray wolf), Capreolus capreolus (Roe deer), Rupicapra rupicapra (Chamois), Wild goat, Hare, Badger, Marten, and others.
AANP is the only place in our country where all three large carnivore species coexist: Brown bear (Ursus arctos), Gray wolf (Canis lupus), and Eurasian lynx (Lynx lynx balcanicus), as well as all three large ungulate species: Chamois (Rupicapra rupicapra), Wild boar (Sus scrofa), and Roe deer (Capreolus capreolus).
Illyrian beech forests (Aremonio-Fagion) in the Gashi Valley are a habitat for a rich fauna. It is important as a habitat for large mammals such as Brown bear (Ursus arctos), Gray wolf (Canis lupus), Wild boar (Sus scrofa), Chamois (Rupicapra rupicapra), and Roe deer (Capreolus capreolus), as well as small mammals.
A significant portion of the park's territory is covered by the habitat of 9150 Medio-European Beech forests on limestone of the Cephalanthero-Fagion alliance and less in the type: 9130 Asperulo-Fagetum Beech forests. Beech habitats in PKN are considered as new forests, not more than 80-100 years old, except for those in Seferce and the Gashi Valley.
According to the Natura 2000 codes the Illyrian Fagus sylvatica forests (Aremonio-Fagion) holds the code 91K0 and from the exisitng literature the habitat covers a surface at about 41.45% of the park.
Habitat 91K0 Illyrian Beech Forests (Aremonio-Fagion) plays an important role in erosion control, as well as being one of the crucial shelters for many species of fauna and flora. Among the species that grow or inhabit beech forests are mentioned: Acer heldreichii, Acer hyrcanum, Betula pendula, Taxus baccata, Helleborus multifidus, Melampyrum doerfleri, Melampyrum heracleoticum.
Thethi, Albanian Alps © Alketa Misja Photography